The apos are centrally attached, subsessile, up to half a mm diameter excluding hairs, light beige.

The marginal hairs are 102-137 × 5.8-7.1 µm, tapering to a point, without crystals, only slighly refractive content, 5-7 times septate, faintly granulose but perhaps not all the way to the very apex. In addition also some shorter hairs less acute, granulose to the apex.

The asci are 34-44 × 3.8-5.5 µm, eight-spored, without croziers, spores biseriate, apex slightly blue with IKI.

The paraphyses are 53-63 × 4.3-5.4 µm, lanceolate, with low-refractive content, protruding 15-25 µm beyond the ascus tips.

The spores are 6.8-9.1 × 1.5-1.8 µm, Q = 3.9-5.4, oil 0.

A few hairs (perhaps 1-2 percent) develop one or a few red brown internal droplets within a few minutes after addition on Lugol.

I have not found many species of neither Dasyscyphella nor Lachnum on ferns in the litterature and not either anything else matching. Perhaps somebody has an idea?

If someone has access to the following paper I would be greatful for a copy:

Bøhler HC (1974) Taxonomical studies on some Norwegian Helotiales (Ascomycetes) on fern remains. Norw J Bot 21:79–100.

Stefan

Hans-Otto Baral, 31-05-2022 07:38

Re : Lachnum(?) on fern

Great find and docu! This must be Lachnaster miniatus (now Psilachnum miniatum).

The protologue also includes rough hairs, unlike the other collections in my folder which is in Psilachnum in Pezizellaceae. What is considered synonymous, Lachnum wasingtonense, has apparently smooth hairs, though not explicitely stated.

This would be worth to be sequenced as I am not at all sure that a Psilachnum may possess warted hairs.

Hans-Otto Baral, 31-05-2022 20:18

Re : Lachnum(?) on fern

It is correct that this species appears to be unable to form the ascus bubble. It has unusual characters, so maybe it is not a real Psilachnum, but what then?